1 0 a b c 0 c1 03_duplication 01_speciation a2 b4_duplication b1 b2 02_speciation a1 b3 0 c1 03_duplication 01_speciation a2 b4_duplication b1 b2 02_speciation a1 b3 0 c1 03_duplication 01_speciation a2 b4_duplication b1 b2 02_speciation a1 b3 0 c1 03_duplication 01_speciation a2 b4_duplication b1 b2 02_speciation a1 b3