1 0 HOSTA BEF. HYBR. a HYBRID HYBRID_ab b HOSTB BEF. HYBR. HYBRID HYBRID_ab a b c HOST2 BEF. HYBR. HOST2 HYBRID HYBRID_H1H2 HOST1 HOST1 BEF. HYBR. HYBRID HYBRID_H1H2 HOST2 HOST1 root symbiot in C DUPLICATION symbiot in HOST2 hybridation symbiote in HOST2 HOST1 hybridation symbiote in HOST2 HOST2 BEF. HYBR.2 HOST2 BEF. HYBR.11 HOST2 BEF. HYBR.12 03_duplication 01_speciation a2 b4_duplication b1 b2 02_speciation HOSTA BEF. HYBR. A1 LOSS b3